A large international research team, directed by the Portuguese archaeologist João Zilhão and including Binghamton University anthropologist Rolf Quam, has found the oldest fossil human cranium in Portugal, marking an important contribution to knowledge of human evolution during the middle Pleistocene in Europe and to the origin of the Neandertals.The cranium represents the westernmost human fossil ever found in Europe during the middle Pleistocene epoch and one of the earliest on this continent to be associated with the Acheulean stone tool industry. In contrast to other fossils from this same time period, many of which are poorly dated or lack a clear archaeological context, the cranium discovered in the cave of Aroeira in Portugal is well-dated to 400,000 years ago and appeared in association with abundant faunal remains and stone tools, including numerous bifaces (handaxes)."This is an interesting new fossil discovery from the Iberian Peninsula, a crucial region for understanding the origin and evolution of the Neandertals," said Quam, an associate professor of anthropology at Binghamton University, State University of New York. "The Aroeira cranium is the oldest human fossil ever found in Portugal and shares some features with other fossils from this same time period in Spain, France and Italy. The Aroeria cranium increases the anatomical diversity in the human fossil record from this time period, suggesting different populations showed somewhat different combinations of features."