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#189658 / #1 | ||
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Pleistocene person
Mod: E&O, S/S, History
Join Date: Mar 2008
Location: London, UK.
Posts: 15,334
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Casey is at it again, this time in response to the latest paper that Per and colleagues have produced. Of course it wouldn't be a Luskin screed without some smug self-aggrandizement; he claims that Boisvert (the lead author of Per's recent paper) admits to an anti-Tiktaalik point he has made before:
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He ends to article with some mental gymnastics concerning Tiktaalik vs Panderichthys. It's all very half arsed, but there we go. http://www.evolutionnews.org/2008/09...tiktaalik.html
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#189723 / #2 | |
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just as bad
Join Date: Mar 2008
Posts: 6,846
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Yeah, so Tiktaalik is crap because Pandrichthys has "more tetrapod-like" fingers, and, at the same time, the distal radials of Panderichthys are too flat and irregular (compared to Tiktaalik's) to even be considered "fingers".
Riiiight. That's Casey Luskin for you. Anyway, my guess was that Per was refering to the arrangement of the distal radials in Panderichthys: A basal line of bones that emegre palallel to each other. The following quote from this article (kindly provided by Rogue06 at TheologyWeb) seems to support that: Quote:
Perhaps that pattern might also make sense, if you consider that, for Tiktaalik, the "digits" were inside the 'fleshy' part of the fin- and were used, more or less, as a "pad" to support the animal when doing its "push-up". In Acanthostega, the bones were used to form the paddle-like structure of the limb, so a fan-like arrangement with single lines of parallel bones would be more beneficial. I guess Per himself can clarify it all for us (I love Talk Rational! )
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If your thinking leads to results like herds of asexual unicellular elephants bobbing around the ocean under a Precambrian moon, it's clear your thinking has gone completely off the rails somewhere. Last edited by Faid; 10-01-2008 at 01:35 PM. |
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#189857 / #3 |
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Senior Member
Join Date: Mar 2008
Posts: 955
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Regarding the "fragments of damaged bone" comment (which may be out of context in any case: I haven't actually seen what Mike Coates said) it's a non-starter. The fin is lying undisturbed in soft sediment, the radials are embedded in an undisturbed and undistorted envelope of scales-plus-fin-rays, and none of the other endoskeletal bones shows any sign of breakage. If that's really what Mike said, he should have known better.
Casey Luskin is not even worth addressing. |
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#189867 / #4 | |||
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Pleistocene person
Mod: E&O, S/S, History
Join Date: Mar 2008
Location: London, UK.
Posts: 15,334
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#195051 / #6 |
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just as bad
Join Date: Mar 2008
Posts: 6,846
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Per, I was wondering about what you were quoted saying, about Tiktaalik being an evolutionary "step backwards" in digit formation. Does that mean that the row of distal radials in Panderichthys led to the creation of "proto-digits" with phalanges, and those digits were later fused in some places, creating the pattern we see in Tiktaalik?
Or is it possible that Tiktaalik's pattern was the first to be produced from Panderichthys' single proximal row, and divisions or duplications later led to the single-line parallel arrangement of digits in Acanthostega? Which is more probable in your opinion? Also, do we have any clues as to how the distal radials in Ventastega were arranged?
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If your thinking leads to results like herds of asexual unicellular elephants bobbing around the ocean under a Precambrian moon, it's clear your thinking has gone completely off the rails somewhere. |
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#195967 / #7 |
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Senior Member
Join Date: Mar 2008
Posts: 955
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Hi Faid,
"Step backwards" is perhaps something of a simplification, but the fin skeleton of Tiktaalik is in certain respects less limb-like than that of Panderichthys. From the shoulder out to the wrist (or from the hip out to the ankle) the tetrapod limb skeleton essentially consists of an axis - one element following another in a straight line - and a series of branches that come off towards the anterior. The humerus or upper arm bone is the first axial element; of the two forearm bones, the ulna (on the "little finger" side of your arm) is the second axial element whereas the radius (on the "thumb" side of your arm) is the first anterior branch. This pattern continues into the wrist, but is most easily seen there in primitive-ish living tetrapods such as salamanders; in ourselves, some of the wrist bones have fused in ways that obscure the pattern. Anyway, the most posterior wrist bone (i.e. just below the little finger) is the ulnare, which is the third axial element. Just anterior to it is the intermedium, which is the second anterior branch. Outside of the wrist bones lie the digits, arranged in a kind of fan shape. We can summarise some of the characteristics of the tetrapod limb skeleton like this: Ulna - long. Usually same length as radius. Ulnare - short. A little wrist bone, much shorter than the ulna. The last element of the axis. Digits - arranged in a fan shape. In Acanthostega (d below), which is a very primitive tetrapod, the radius is still longer than the ulna but all the other characteristics seem to be there. Now, in Panderichthys, we find that the ulnare is much shorter than the ulna and really looks like a wrist bone. Furthermore, the ulnare is the last axial element, and beyond it the distal radials are arranged in something of a fan shape. But in Tiktaalik the ulna and ulnare are equal in size, there are two more axial elements beyond the ulnare, and the distal radials are arranged bipinnately (i.e. like the leaflets of a palm leaf) on either side of this distal axis. In all these respects Tiktaalik's fin skeleton (c) is less limb-like than that of Panderichthys (b) and compares more closely with lobe-finned fishes (e.g. Eusthenopteron, a): ![]() The interesting question is whether this means that: The detailed similarities between Panderichthys and tetrapods are convergent, or - The seemingly more primitive fin skeleton of Tiktaalik represents an evolutionary reversal, or - The current phylogenetic hypothesis is wrong and Panderichthys is actually more closely related to tetrapods than Tiktaalik. One of these three explanations must be correct, but it is not yet possible to tell which one. Sadly, we don't have any data at all on the limb structure of Ventastega. A reasonable guess is that they resembled those of Acanthostega, because the limb girdles are similar, but no limb bones have been found. |
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#196871 / #8 |
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Pleistocene person
Mod: E&O, S/S, History
Join Date: Mar 2008
Location: London, UK.
Posts: 15,334
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Regular Pandas Thumb creationist FL has been happily spouting forth about this latest research, starting from here:
http://pandasthumb.org/archives/2008...comment-169203 I have invited him to Talk Rational to discuss these issues. He claims he will post this weekend (I'll believe it when I see it). In advance, he relies heavily on this article on Tiktaalik: http://www.earthhistory.org.uk/techn...taalik-roseae/ and this more general one on tetrapod evolution: http://www.earthhistory.org.uk/trans...-to-amphibian/ If you've got time at the moment Per, I'd appreciate your thoughts on these.
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#199053 / #9 |
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It's Elementary
Administrator
Join Date: Mar 2008
Posts: 18,074
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I'll ask the awkward question (I already have some possible thoughts, so I want to see if I am right) - how might you be able to tell? You imply that it may be possible in future...
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The Feynmann Algorithm: (1) Write down the problem (2) Think real hard (3) Write down the solution
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#199510 / #12 | |
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Senior Member
Join Date: Mar 2008
Posts: 955
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#205071 / #13 |
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Junior Member
Join Date: Oct 2008
Posts: 4
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Well, I'm here. And thanks for inviting me, SteveF, it's a nice cozy place you got here.
I use the handle Mellotron for this forum, but I'm also the "FL" that SteveF spoke of. I have no intention of doing any long-drawn-out debating in this thread, and I know that Dr. Ahlberg's time is restricted and valuable, but I was asked to present some arguments to Dr. Ahlberg for his commentary. So I'll just ask about a couple of items a little later today, and that should be all. (For this thread, anyway.... heh-heh-heh.) Mellotron ![]() |
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#206019 / #14 |
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Senior Member
Join Date: Mar 2008
Posts: 955
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Hi Mellotron, and welcome to TalkRational!
As luck would have it I have just started my main teaching block for the autumn (I'm lecturing 9 am to 2 pm today - hooray!!) but I'll try to find time to answer your points. Cheers, Per |
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#213408 / #15 | ||||
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Junior Member
Join Date: Oct 2008
Posts: 4
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No problem! And my apologies for my own delay in getting back to this forum.
At best, I'll only be asking a couple questions. The first one is simply, do you agree with Casey Luskin's main observation regarding Tiktaalik? Quote:
As I await your answer to that question, I'll go ahead and share mine. I agree with Luskin's statement. I think his main observation is correct. Here's a couple examples of the hype Luskin spoke of, items that I googled on my own: Quote:
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These examples lead me to conclude that indeed the Tiktaalik hype seriously does NOT match the actual reality, and thus Luskin is therefore correct in his main observation. So, back to the question: Do you agree or disagree with Luskin's main observation? **** There is a second question based on a snippet from the creationist website Earth History A New Approach (this is separate and different from Casey Luskin's Evolution News & Views article), and I'll place that question on the table now. Just asking to see if you agree or disagree, and why. In an April 12, 2008 article, "Tiktaalike roseae -- a missing link?", the Earth History website wrote: Quote:
Do you agree or disagree that Tiktaalik failed that prediction? Thanks again for your time. Mellotron ![]() |
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#213567 / #16 |
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just as bad
Join Date: Mar 2008
Posts: 6,846
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Um, I think we should wait for Per to respond, so I'll refrain from commenting, even though I'll have to fight the urge...
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If your thinking leads to results like herds of asexual unicellular elephants bobbing around the ocean under a Precambrian moon, it's clear your thinking has gone completely off the rails somewhere. |
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#213871 / #17 | |||||
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The Witchdoctor
Join Date: Apr 2008
Location: MO
Posts: 2,257
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Quote:
http://www.youtube.com/watch?v=uiccF7DDUQc Elká, waiting, chin in hand.
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Never miss a good chance to shut up. Will Rogers The correct answer is, None of the above. |
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#213924 / #18 |
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Senior Member
Join Date: Mar 2008
Posts: 955
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Hi Mellotron,
That's a pretty relevant question you're asking, and I am going to give the best and most nuanced answer I can. Please pay careful attention to every part of the argument. Is there a problem with the way Tiktaalik has been hyped? Yes, there is. Does this mean that it has been "dethroned" by Panderichthys, or that it is no longer relevant to the question of tetrapod origins? No it doesn't. The underlying issue is that research into major evolutionary transitions (from fish to tetrapod, from reptile to bird, that kind of thing) does not and should not focus on Unique Missing Links, but is frequently portrayed that way in the media. Nor is this a straightforward case of the wicked sensationalist media distorting the views of the innocent scientists: there is always a temptation for the scientists to boost the profile of their particular find, and in the case of Tiktaalik I do feel that the authors have not always been careful enough to ensure that the animal was placed in its proper context by news reporters. By allowing it to be portrayed as a unique missing link devoid of context, standing all alone in the beam of the spotlight as it were, they have inadvertently obscured some of its real significance. Note, however, that I am speaking here of popular media reports written by others, not the actual scientific papers by the discoverers. Two things need to be understood in relation to the study of major transitions: the first is that these are not simple two-step affairs from a starting point via a "missing link" to an end point ("normal" fish -> Tiktaalik -> fully developed land vertebrate) but rather very gradual processes involving lots of steps, and the second is that it is extremely difficult to identify direct ancestor-descendant sequences in the fossil record. The first of these points is I think intuitively obvious, but the second needs a bit of consideration. In essence it boils down to the fact that we have in the fossil record, in all likelihood, only a very small sample of the animals that lived at any one time, and the time sequence is full of gaps as well. With very recent fossils such as those from the Ice Age, where the time sampling is densest, we can be reasonably confident that we are retrieving some genuine ancestor-descendant chains: for example, it really does seem to be the case that Mammuthus primigenius (the woolly mammoth) is descended from Mammuthus trogontherii, which in turn is descended from Mammuthus meriodinalis, and that Homo sapiens is directly or indirectly descended from Homo erectus. However, with fossils from the Devonian we don't have much hope for that kind of resolution, so we don't look for direct ancestors but relatives. The characteristics shared by the different fossil forms that we find, with each other and with now living groups, tell us about how they are related to each other and what their common ancestors must have been like. Think about it this way: Consider a person you know, that you are not related to - a colleague at work or whatever. You belong to the same species and have numerous features in common, both morphological (people all basically look the same apart from gender differences, only the incidentals differ) and molecular (all people have very similar genes). This means that you actually are related, despite having no known family relationship; somewhere back in time, maybe thousands of years ago, the two of you share a last common ancestor. Now here's the key point: you have virtually no hope of ever finding the bones of that ancestor, but you can actually say quite a lot about him/her because every attribute shared by the two of you (Presence of nose? Check! Presence of the gene Hoxd13? Check! Etc. etc.) must also have been there in your common ancestor. Do you see how it works? Now we can return to Tiktaalik, Panderichthys and their kin. The real story here is that there exists a set of fossils that are more similar to the extant Tetrapoda (land vertebrates) than to any other extant vertebrate group, but which nevertheless do not have the full set of land vertebrate characteristics. We hypothesise that these fossils belong to the lineage that gave rise to land vertebrates, and refer to them collectively as the tetrapod stem group. Some members of the stem group, such as Eusthenopteron, are pretty conventional-looking fishes that have a few anatomical features in common with land vertebrates (for example internal nostrils and a humerus-radius-ulna limb skeleton inside the pectoral fin) but show no actual adaptations for life on land. These would appear to form the lower part of the tetrapod stem group. Other members of the stem group such as Ichthyostega and Acanthostega have all those same land vertebrate characteristics plus additional ones like limbs with toes, and evidently had some degree of terrestrial capability, but retain a few fish characteristics (for example a real tail fin with fin rays) that are not seen in any extant tetrapods. These appear to form the upper part of the stem group. So, from this information we can infer the existence of two common ancestors: one very ancient common ancestor of Eusthenopteron (and other similar fishes), Acanthostega + Ichthyostega and extant tetrapods, and another somewhat more recent common ancestor of just Acanthostega + Ichthyostega and extant tetrapods. The second of these common ancestors must be a descendant of the first. Furthermore, we can infer that the first of these common ancestors possessed internal nostrils and a humerus-radius-ulna complex in the pectoral fin, while the second ancestor also possessed these characteristics plus digits, a pelvis attached to the backbone, etc.etc. This begins to give us a breakdown of the evolutionary transformation from fish to tetrapod. Panderichthys and Tiktaalik both belong in the middle part of the stem group, somewhere between Eusthenopteron and Acanthostega. In most respects, Tiktaalik is marginally more tetrapod-like than Panderichthys: for example, it lacks a bony gill cover, which is still present in Panderichthys. For this reason the original discoverers argued that Tiktaalik occupies a somewhat higher position in the stem group. In most respects, P. and T. are very similar. At a minimum (i.e. without attempting for the moment to resolve exactly how they are related to each other), they allow us to infer one new common ancestor in our scheme: the last common ancestor of Panderichthys, Tiktaalik, Acanthostega + Ichthyostega and extant tetrapods. This common ancestor will lie between the two previously discussed ones in the stem group, being a descendant of the first common ancestor and an ancestor of the second. This new common ancestor shows us an intermediate step in the transformation: it has, for example, already lost the dorsal and anal fins (unlike Eusthenopteron which retains these fins) but has not yet acquired digits (unlike Acanthostega, Ichthyostega and extant tetrapods). If we accept the hypothesis that Tiktaalik goes above Panderichthys in the stem group, things get a bit more interesting because the new common ancestor that I just described splits into two: a slightly older common ancestor of Panderichthys, Tiktaalik, Acanthostega + Ichthyostega and extant tetrapods, and a slightly younger common ancestor of just of Tiktaalik, Acanthostega + Ichthyostega and extant tetrapods. These two common ancestors will be very similar, and this is their value: they begin to define in great detail the small evolutionary steps occurring in this part of the tree. You should now be able to see that the importance of Tiktaalik lies very largely in its context, and at least partly in the fact that is is not very different from Panderichthys. Together, all these fossil forms help to map out the transition from fish to land vertebrate that occurred in the tetrapod stem group. So what about the pectoral fin business? Well, the pectoral fin skeleton of Tiktaalik is in many ways a good prototype for a tetrapod forelimb skeleton. It has the same basic humerus-radius-ulna architecture, and the shape of the humerus in particular is very like that of a tetrapod. The humerus and radius of Panderichthys compare closely with those of Tiktaalik. Surprisingly, however, the pectoral fin skeleton proves to be somewhat more limb-like than that of Tiktaalik in having a lung ulna, short ulnare (a wrist bone), and distal radials (digit equivalents) that are arranged in a transverse row rather like toes. In Tiktaalik the ulna and ulnare are about the same size and the distal radials are arranged differently. What does this mean? Well, it obviously clashes with the other character distributions (gill cover etc.) that place Tiktaalik closer to tetrapods than Panderichthys, so three interpretations appear plausible: either 1) the similarities between the fin skeleton of Panderichthys and the limb skeletons of tetrapods are due to parallel evolution, or 2) the non-limb-like characteristics of the fin skeleton of Tiktaalik are just unique specialisations of that animal, or 3) the currently accepted phylogeny is wrong and Panderichthys is in fact closer to tetrapods than Tiktaalik. Personally I believe that 1) and 2) are more probable than 3), but we are not yet in a position to settle this question - and may never be. However, this has only a minor negative impact on our understanding of tetrapod origins. As regards the pelvis of Tiktaalik, the situation is somewhat different. The pelvis has not been described, so Luskin is either seriously confused or dishonest in making claims about it. There is no basis for claiming that Tiktaalik "was more of a ‘front-wheel drive’ animal than Panderichthys was". However, if we assume for the moment that Tiktaalik will prove to have exactly the same fish-like kind of pelvis that we see in Panderichthys, this still does not mean that Tiktaalik fails any sort of prediction. The reason is simple: your prediction assumes, invalidly, that any fossil falling between Panderichthys and Acanthostega on the tree will be neatly intermediate between them in every respect. There is no reason to believe that that will be the case. You are basically arguing that, because 5 is intermediate between 2 and 9, any randomly picked number that is not 5 is not an intermediate between 2 and 9. I trust you can see the fallacy. |
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#213951 / #19 |
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Science n00b
Join Date: Apr 2008
Location: Adelaide, South Australia
Posts: 1,822
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Per, he's talking about panderichthys' pelvis for the second question isn't he, not Tiktaalik's.?
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"There probably is no teapot. Now stop worrying and enjoy your life." Humber: You say that Goodman's cart travels faster than windspeed, so how do you explain that adding mass to that cart, will stop that happening? |
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#213961 / #21 | ||
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не могу устоять
Join Date: Apr 2008
Location: UK
Posts: 1,652
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#213962 / #22 |
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Science n00b
Join Date: Apr 2008
Location: Adelaide, South Australia
Posts: 1,822
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Ahh yep, sorry. I misread his question. It's been a long day.
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"There probably is no teapot. Now stop worrying and enjoy your life." Humber: You say that Goodman's cart travels faster than windspeed, so how do you explain that adding mass to that cart, will stop that happening? |
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#213993 / #23 |
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A fuzzy, winged spork.
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I'm betting the creationist crap site that he got his unformation (intentional spelling) from probably got it from the site that Martin B dissected long ago. But we all know how creationists won't let little things like facts and reality get in the way of spreading their crap.
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#214007 / #24 |
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Science n00b
Join Date: Apr 2008
Location: Adelaide, South Australia
Posts: 1,822
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"unformation" - I love it.
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"There probably is no teapot. Now stop worrying and enjoy your life." Humber: You say that Goodman's cart travels faster than windspeed, so how do you explain that adding mass to that cart, will stop that happening? |
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#216291 / #25 | ||
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Junior Member
Join Date: Oct 2008
Posts: 4
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Meanwhile, sincere thanks to all who responded, and particularly Dr. Ahlberg. Dr. Ahlberg, I have seen that "context" argument once or twice before (but not in as much detail and honesty as your explanation), and it seems clear that the "context" line of argument will be the direction in which evolutionists continue to maintain that Tiktaalik is part of the fish-to-tetrapod transition. But again, I appreciate your detailed explanation and "nuances", and will save it for my own study, to compare and contrast it with the Luskin and Earth History articles. ****** One other thing. You said..... Quote:
In three different discussion forums, you're the ONLY evolutionist I've met who has displayed such honesty and candor about the situation. (Just want to say a sincere thanks for that. Most refreshing.) So, like I said, I don't have any plans for long-drawn-out debate here. It's enough for me to put forth the two questions and to receive your receive your response. Thanks once again. Mellotron Last edited by Mellotron; 10-21-2008 at 05:57 PM. |
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